Research Interests: Animal ecological physiology; energetic costs of reproduction and embryonic development; sexual dimorphism; herpetology.

Most of my recent work has focused on internal anatomical and physiological dimorphisms related to reproductive cost in spring breeding frogs and salamanders.  I am also interested in  the influence of environmental quality within amniote eggs on hatchling phenotype (turtles and birds), water balance and desiccation tolerance in crayfish, correlates between morphology and interspecific competition in crayfish, the impact of trematode parasitism on fecundity and stored energy in amphibians, and the effects of thermal pollution on native mussel species.